Mérida Sunangel
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The Mérida sunangel (''Heliangelus spencei'') is a species of
hummingbird Hummingbirds are birds native to the Americas and comprise the biological family Trochilidae. With about 361 species and 113 genera, they occur from Alaska to Tierra del Fuego, but the vast majority of the species are found in the tropics aro ...
in the "coquettes", tribe
Lesbiini Lesbiini is one of the two tribes that make up the subfamily Lesbiinae in the hummingbird family Trochilidae. The other tribe is Heliantheini (brilliants). The informal name "coquettes" has been proposed for this group as the largest genus, ''L ...
of subfamily
Lesbiinae Lesbiinae is one of the six subfamilies that make up the hummingbird family Trochilidae. The subfamily is divided into two tribes: Heliantheini ("brilliants") containing 14 genera and Lesbiini ("coquettes") containing 18 genera. Phylogeny A mo ...
. It is endemic to
Venezuela Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in th ...
.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved May 27, 2021


Taxonomy and systematics

The taxonomy of the Merida sunangel is not settled. The
International Ornithological Committee The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...
(IOC), the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 2022 ...
, and
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding ...
's
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. T ...
(HBW) treat it as a species.Clements, J. F., T. S. Schulenberg, M. J. Iliff, T. A. Fredericks, J. A. Gerbracht, D. Lepage, S. M. Billerman, B. L. Sullivan, and C. L. Wood. 2022. The eBird/Clements checklist of birds of the world: v2022. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ retrieved November 10, 2022 The South American Classification Committee (SACC) of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its m ...
treats it as a subspecies of the
amethyst-throated sunangel The amethyst-throated sunangel (''Heliangelus amethysticollis'') is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is found in Bolivia, Ecuador, and Peru.HBW and BirdLife International (2020) ''Handbook ...
(''H. amethysticollis'').Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 July 2022. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved July 24, 2022


Description

The Merida sunangel is long. It has a straight black bill. The adult male has a pale blue frontlet just above the bill, a dark green crown, and shining green upperparts. The face has black ear
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which, as the name implies, cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are s ...
. Its throat and upper breast
gorget A gorget , from the French ' meaning throat, was a band of linen wrapped around a woman's neck and head in the medieval period or the lower part of a simple chaperon hood. The term later described a steel or leather collar to protect the th ...
is glittering purple with pinkish scales and has a white pectoral band below it. The rest of the underparts are yellowish to pale buff with round green spots. The central tail feathers are dark green and the outer feathers blackish, occasionally with pale tips. The adult female has less black on the face. Its throat is maroon with white and green scaling. Its underparts have fewer green spots than the male's and appears more yellowish. Juveniles are thought to resemble the female.del Hoyo, J., I. Heynen, N. Collar, P. F. D. Boesman, G. M. Kirwan, and C. J. Sharpe (2020). Amethyst-throated Sunangel (''Heliangelus amethysticollis''), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.amtsun1.01 retrieved January 20, 2022


Distribution and habitat

The Merida sunangel is found only in the
Andes The Andes, Andes Mountains or Andean Mountains (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long, wide (widest between 18°S – 20°S ...
of Mérida state in northwestern Venezuela. It inhabits rather open landscapes such as shrubby forest edges, openings within forest, and brushy pastures. In elevation it ranges between .


Behavior


Movement

The Merida sunangel is sedentary.


Feeding

Little is known about the Merida sunangel's feeding habits and diet, but they are believed to essentially the same as those of its close relative,
Longuemare's sunangel Longuemare's sunangel (''Heliangelus clarisse'') is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is found in Colombia and Venezuela.HBW and BirdLife International (2020) ''Handbook of the Birds of the ...
(''H. clarisse''). That species feeds on nectar and also includes insects in its diet. It defends feeding territories that are dense with nectar sources. It typically feeds at heights up to , often along streams. It captures insects by
hawking Hawking may refer to: People * Stephen Hawking (1942–2018), English theoretical physicist and cosmologist * Hawking (surname), a family name (including a list of other persons with the name) Film * ''Hawking'' (2004 film), about Stephen Ha ...
from a perch.


Breeding

The only known Merida sunangel nest was found in June. It was a downy cup perched on a small root under an overhang in a road cut. It held two white eggs. Nothing else is known about the species' breeding
phenology Phenology is the study of periodic events in biological life cycles and how these are influenced by seasonality, seasonal and interannual variations in climate, as well as environmental factor, habitat factors (such as elevation). Examples includ ...
.


Vocalization

The Merida sunangel's call is "a repeated, high-pitched, cricket-like, short trill" that is given both while perched and in flight.


Status

The
IUCN The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed the Merida sunangel as being of Least Concern, though its population size and trend are not known. It has a small range but about 3/5 of it is within protected areas. The principal threats are "expansion of agricultural frontier and habitat fragmentation."


References

{{Taxonbar, from=Q1957721
Mérida sunangel The Mérida sunangel (''Heliangelus spencei'') is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is endemic to Venezuela.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and Bird ...
Birds of the Venezuelan Andes
Mérida sunangel The Mérida sunangel (''Heliangelus spencei'') is a species of hummingbird in the "coquettes", tribe Lesbiini of subfamily Lesbiinae. It is endemic to Venezuela.HBW and BirdLife International (2020) ''Handbook of the Birds of the World and Bird ...